From left to right, Philo, Cioki, Marcel, and Zawadi. All are children of the Mwangan Empress Senna, who is a tigon, and the Shupavuan Kipande, who is a second generation leopon. The cubs were born out of physical desire (and one out of a godly wish), rather than love. 

In fact, the Empress Senna holds custody of Philo, Cioki, and Marcel, all of whom were born to her, and Kipande has custody of Zawadi, a child who materialized due to a wish.

Truthfully, Cioki only shares half her blood with her brothers. Her father is none other than the leopard Tibor.

All cubs are what we call "Turbo Hybrids", or a hybrid with multiple species. Due to the nature of their birth, none can ever have children of their own. 

Philo and Marcel were designed by me, Zawadi and Cioki were designed by Kit.

Philo belongs to Yoki
Marcel belongs to Mx. Corvidae
Zawadi belongs to Kit
Cioki belongs to me.